William G. Eberhard

218 publications found


Aisenberg, Anita, Barrantes, Gilbert and Eberhard, William G. 2014. Hairy kisses: tactile cheliceral courtship affects female mating decisions in Leucauge mariana (Araneae, Tetragnathidae). Doi: 10.1007/s00265-014-1844-2 [Download]
Méndez, Vivian and Eberhard, William G. 2014. Removal of genital plugs and insemination by males with normal and experimentally modified palps in Leucauge mariana (Araneae: Tetragnathidae). Journal of Arachnology, 42(2):284-292 [Download]


Ang, Yuchen; Puniamoorthy, Jayanthi; Pont, Adrian C.; Bartak, Miroslav; Blanckenhorn, Wolf U.; Eberhard, William G.; Puniamoorthy, Nalini; Silva, Vera C.; Munari, Lorenzo; Meier, Rudolf. 2013. A plea for digital reference collections and other science-based digitization initiatives in taxonomy: Sepsidnet as exemplar. Systematic Entomology, 38(3): 637-644. [Download]


Barrantes, Gilbert; Eberhard, William G.. 2012. Extreme Behavioral Adjustments by an Orb-Web Spider to Restricted Spaces. Ethology, 118(5): 438-449.. [Download]
Briceno, R. D.; Eberhard, William G.. 2012. Spiders avoid sticking to their webs: clever leg movements, branched drip-tip setae, and anti-adhesive surfaces. Naturwissenschaften, 99(4): 337-341.. [Download]
Eberhard, William G.; Hesselberg, Thomas. 2012. Cues that Spiders (Araneae: Araneidae, Tetragnathidae) Use to Build Orbs: Lapses in Attention to One Set of Cues because of Dissonance with Others?. Ethology, 118(7): 610-620. [Download]
Eberhard, William G.; Wcislo, William T.. 2012. Plenty of room at the bottom? Tiny animals solve problems of housing and maintaining oversized brains, shedding new light on nervous-system evolution. American Scientist, 100(3): 226-233.. [Download]


Briceño, R. D. & Eberhard, W. G. 2011. The hub as a launching platform: rapid movements of the spider Leucauge mariana (Araneae: Tetragnathidae) as it turns to attack prey. Journal of Arachnology, 39(1), 102–112. [Download]
Eberhard, W. G. & Wcislo, W. T. 2011. Morphological and behavioral consequences of physiological limitations in miniature arthropods. Advances in Insect Physiology. [Download]
Eberhard, W. G. 2011. Are smaller animals behaviourally limited? Lack of clear constraints in miniature spiders. Animal Behaviour, 81(4), 813-823. [Download]
Eberhard, W. G. 2011. Experiments with genitalia: a commentary. Trends in Ecology and Evolution 26(1), 17-21. doi:10.1016/j.tree.2010.10.009 [Download]
Quesada, Rosannette; Triana, Emilia; Vargas, Gloria; Douglass, John K.; Seid, Marc A.; Niven, Jeremy E.; Eberhard, William G.; Wcislo, William T.. 2011. The allometry of CNS size and consequences of miniaturization in orb-weaving and cleptoparasitic spiders. Arthropod Structure & Development, 40(6): 521-529.. [Download]


Barrantes, G. & Eberhard, W. G. 2010. Ontogeny repeats phylogeny in Steatoda and Latrodectus spiders. Journal of Arachnology, 38(3), 485–494. [Download]
Briceño, R. D., Wegrzynek, D., Chinea-Cano, E., Eberhard, W. G. & dos Santos Rolo, T. 2010. Movements and morphology under sexual selection: tsetse fly genitalia. Ethology Ecology & Evolution, 22(4), 385–391. [Download]
Eberhard, W. G. & Huber, B. A. 2010. Spider Genitalia. Pp. 249-284 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals. [Download]
Eberhard, W. G. 2010. New types of behavioral manipulation of host spiders by a parasitoid wasp. Psyche, Article ID 950614, 1-4. [Download]
Eberhard, W. G. 2010. Possible functional significance of spigot placement on the spinnerets of spiders. Journal of Arachnology, 38(3), 407–414. [Download]
Eberhard, W. G. 2010. Rapid divergent evolution of genitalia. Pp. 40-78 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals. [Download]
Eberhard, W. G. 2010. Recovery of spiders from the effects of parasitic wasps: implications for fine-tuned mechanisms of manipulation. Animal Behaviour, 79(2), 375–383. [Download]
Eberhard, W.G. (2010) Cryptic Female Choice. In: Breed M.D. and Moore J., (eds.) Encyclopedia of Animal Behavior, volume 1, pp. 430-434 Oxford: Academic Press. [Download]
Gonzaga, M., Sobczak, J., Penteado-Dias, A. & Eberhard, W. 2010. Modification of Nephila clavipes (Araneae Nephilidae) webs induced by the parasitoids Hymenoepimecis bicolor and H. robertsae (Hymenoptera Ichneumonidae). Ethology Ecology & Evolution, 22(2), 151–165. [Download]
Moya, J., R. Quesada, G. Barrantes, WG Eberhard, I. Escalante, C. Esquivel, A. Rojas, E. Triana and A. Arias. 2010. Egg sac construction by folding dead leaves in Pozonia nigroventris and Micrathena sp. (Araneae: Araneidae). Journal of Arachnology, 38(2):371-373. [Download]


Aisenberg, A. & Eberhard, W. G. 2009. Female cooperation in plug formation in a spider: effects of male copulatory courtship. Behavioral Ecology, 20(6), 1236. [Download]
Briceño, R. D. & Eberhard, W. G. 2009. Experimental demonstration of possible cryptic female choice on male tsetse fly genitalia. Journal of Insect Physiology, 55(11), 989–996. [Download]
Briceño, R. D. & Eberhard, W. G. 2009. Experimental modifications imply a stimulatory function for male tsetse fly genitalia, supporting cryptic female choice theory. Journal of Evolutionary Biology, 22(7), 1516–1525. [Download]
Eberhard, W. G. & Gelhaus, J. K. 2009. Genitalic stridulation during copulation in a species of crane fly, Tipula (Bellardina) sp. (Diptera: Tipulidae). Revista de Biología Tropical, 57(Suppl. 1), 251–256. [Download]
Eberhard, W. G. 2009. Book Review: Art for art’s sake. Ethology, 115, 1-2. [Download]
Eberhard, W. G. 2009. Postcopulatory sexual selection: Darwin’s omission and its consequences. Proceedings of the National Academy of Sciences, 106(Suppl 1), 10025-10032. [Download]
Eberhard, W. G. 2009. Static allometry and animal genitalia. Evolution, 63(1), 48–66. [Download]
Eberhard, W. G., Rodriguez, R. L. & Polihronakis, M. 2009. Pitfalls in understanding the functional significance of genital allometry. Journal of Evolutionary Biology, 22(3), 435–445. [Download]
Peretti, A. V. & Eberhard, W. G. 2009. Cryptic female choice via sperm dumping favours male copulatory courtship in a spider. Journal of Evolutionary Biology, 23(2), 271–281. [Download]
William G. Eberhard. 2009. Evolution of genitalia – theories, evidence, and new directions. Genetica. 2009. [Download]


Barrantes, G., Eberhard, W. G. & Weng, J. L. 2008. Seasonal patterns of parasitism of the tropical spiders Theridion evexum (Araneae, Theridiidae) and Allocyclosa bifurca (Araneae, Araneidae) by the wasps Zatypota petronae and Polysphincta gutfreundi (Hymenoptera, Ichneumonidae). Revista de Biología Tropical, 56(2), 749–754. [Download]
Eberhard, W. G. 2008. Araneus expletus (Araneae, Araneidae): another stabilimentum that does not function to attract prey. Journal of Arachnology, 36(1), 191–194. [Download]
Eberhard, W. G., Agnarsson, I. & Levi, H. W. 2008. Web forms and the phylogeny of theridiid spiders (Araneae: Theridiidae): chaos from order. Systematics and Biodiversity, 6(4), 415–475. [Download]
Eberhard, W. G., Barrantes, G. & Madrigal-Brenes, R. 2008. Vestiges of an orb-weaving ancestor? The “biogenetic law” and ontogenetic changes in the webs and building behavior of the black widow spider Latrodectus geometricus (Araneae Theridiidae). Ethology Ecology & Evolution, 20(3), 211–244. [Download]


Baena, ML; Eberhard, W. G. 2007. Appearances deceive: female "resistance" behaviour in a sepsid fly is not a test of male ability to hold on. Ethology Ecology & Evolution. 19 (1) pp. 27-50. [Download]
Barrantes, G. & Eberhard, W. G. 2007. The evolution of prey-wrapping behaviour in spiders. Journal of Natural History, 41(25-28), 1631–1658. [Download]
Briceño, R. D., Eberhard, W. G. & Robinson, A. S. 2007. Copulation behaviour of Glossina pallidipes (Diptera: Muscidae) outside and inside the female, with a discussion of genitalic evolution. Bulletin of Entomological Research, 97(5), 471–488. [Download]
Briceño, R. D., Eberhard, W. G. & Shelly, T. 2007. Male courtship behavior in Ceratitis capitata (Diptera: Tephritidae) that have received aromatherapy with ginger root oil. Florida Entomologist, 90(1), 175–179. [Download]
Briceño, R. D., Eberhard, W. G., Vilardi, J., Cayol, J. P. & Shelly, T. 2007. Courtship behavior of different wild strains of Ceratitis capitata (Diptera: Tephritidae). Florida Entomologist, 90(1), 15–18. [Download]
Eberhard, W. G. 2007. Miniaturized orb-weaving spiders: behavioural precision is not limited by small size. Proceedings of the Royal Society B: Biological Sciences, 274(1622), 2203-2209. [Download]
Eberhard, W. G. 2007. Stabilimenta of Philoponella vicina (Araneae: Uloboridae) and Gasteracantha cancriformis (Araneae: Araneidae): evidence against a prey attractant function. Biotropica, 39(2), 216–220. [Download]
Eberhard, W. G. 2007. Swarming behavior of two parasitic wasps, Leurus caeruliventris (Hymenoptera: Ichneumonidae) and Conura sp.(Hymenoptera: Chalcididae), at the same site. Journal of the Kansas Entomological Society, 80(3), 255–258. [Download]
Jörger, K. M. & Eberhard, W. G. 2006. Web construction and modification by Achaearanea tesselata (Araneae, Theridiidae). Journal of Arachnology, 34(3), 511–523. [Download]


Daniel Briceño, William Eberhard, Juan Vilardi, Jean-Pierre Cayol and Todd Shelly. 2006. Courtship Behavior of Different Wild Strains of Ceratitis Capitata (Diptera: Tephritidae). [Download]
Eberhard, W. G. 2006. Dispersal by Ummidia spiderlings (Araneae, Ctenizidae): ancient roots of aerial webs and orientation? Journal of Arachnology, 34(1), 254–257. [Download]
Eberhard, W. G. 2006. Sexually antagonistic coevolution in insects is associated with only limited morphological diversity. Journal of Evolutionary Biology. 19(3):657-681. [Download]
Eberhard, W. G. 2006. Sexually Reversed Copulatory courtship roles and possible nuptial feeding in the soldier beetle Ditemnus acantholabus (Champion 1915) (Coleoptera: Cantharidae). Journal of the Kansas Entomological Society, 79(1), 13–22. [Download]
Eberhard, W. G., Barrantes, G. & Weng, J. L. 2006. Tie them up tight: wrapping by Philoponella vicina spiders breaks, compresses and sometimes kills their prey. Naturwissenschaften 93(5), 251-254. [Download]
Flowers, W. & Eberhard, W. G. 2006. Fitting together: copulatory linking in some Neotropical Chrysomeloidea. Revista de Biología Tropical, 54(3), 829–842 [Download]
Jaffé, R., Eberhard, W. G., De Angelo, C., Eusse, D., Gutierrez, A., Quijas, S. & Rodríguez, A. 2006. Caution, webs in the way! Possible functions of silk stabilimenta in Gasteracantha cancriformis (Araneae, Araneidae). Journal of Arachnology, 34, 448–455 [Download]
Peretti, A., Eberhard, W. G. & Briceño, R. D. 2006. Copulatory dialogue: female spiders sing during copulation to influence male genitalic movements. Animal Behaviour, 72(2), 413–421. [Download]
Weng, J. L., Barrantes, G. & Eberhard, W. G. 2006. Feeding by Philoponella vicina (Araneae, Uloboridae) and how uloborid spiders lost their venom glands. Canadian Journal of Zoology, 84(12), 1752–1762. [Download]


Briceño, R. D., Eberhard, W. G. & Quilici, S. 2005. Comparative allometry and sexual behavior of four fruit fly species in the tribe Ceratitidini (Diptera: Tephritidae). Journal of the Kansas Entomological Society, 78(1), 20–33. [Download]
Cordero, C. & Eberhard, W. G. 2005. Interaction between sexually antagonistic selection and mate choice in the evolution of female responses to male traits. Evolutionary Ecology, 19(2), 111–122. [Download]
Eberhard W. 2005. Why study spider sex: special traits of spider facilitate study of sperm competition and cryptic female choice. J. Arachnol. 32:545-556 [Download]
Eberhard, W. G. 2005. Evolutionary conflicts of interest: are female sexual decisions different? American Naturalist, 165(5), S19–S25. [Download]
Eberhard, W. G. 2005. Sexual morphology of male Sepsis cynipsea (Diptera: Sepsidae): lack of support for lock-and-key and sexually antagonistic morphological coevolution hypotheses. The Canadian Entomologist, 137(5), 551–565. [Download]
Eberhard, W. G. 2005. Threading a needle with reinforced thread: intromission in Ceratitis capitata (Diptera, Tephritidae). The Canadian Entomologist, 137(2), 174–181. [Download]


Eberhard, W. G. & Ramirez, N. 2004. Functional morphology of the male genitalia of four species of Drosophila: failure to confirm both lock and key and male-female conflict predictions. Annals of the Entomological Society of America, 97(5), 1007–1017. [Download]
Eberhard, W. G. 2004. Apparent stimulatory function of species-specific male genitalic setae in Aelurus septentrionalis (Hymenoptera: Tiphiidae). Journal of the Kansas Entomological Society, 77(4), 837–849. [Download]
Eberhard, W. G. 2004. Male–female conflict and genitalia: failure to confirm predictions in insects and spiders. Biological Reviews, 79(1), 121–186. [Download]
Eberhard, W. G. 2004. Rapid divergent evolution of sexual morphology: comparative tests of antagonistic coevolution and traditional female choice. Evolution, 58(9), 1947–1970. [Download]
Eberhard, W. G. 2004. Why study spider sex: special traits of spiders facilitate studies of sperm competition and cryptic female choice. Journal of Arachnology, 32(3), 545–556. [Download]
Rodriguez, V., Windsor, D. & Eberhard, W. G. 2004. Tortoise beetle genitalia and demonstrations of a sexually selected advantage for flagellum length in Chelymorpha alternans (Chrysomelidae, Cassidini, Stolaini). Pp. 739-748 in P. Jolivet, J. A. Santiago-Blay & M. Schmitt New developments in the biology of Chrysomelidae. The Hague, The Netherlands: SPB Academic Publications. [Download]


Cordero, C. & Eberhard, W. G. 2003. Female choice of sexually antagonistic male adaptations: a critical review of some current research. Journal of Evolutionary Biology, 16(1), 1–6. [Download]
Eberhard, W. G. & Cordero, C. 2003. Sexual conflict and female choice. Trends in Ecology and Evolution, 18(9), 438–439. [Download]
Eberhard, W. G. 2003. Sexual behavior and morphology of Themira minor (Diptera: Sepsidae) males and the evolution of male sternal lobes and genitalic surstyli. The Canadian Entomologist, 135(4), 569–581. [Download]
Eberhard, W. G. 2003. Substitution of silk stabilimenta for egg sacs by Allocyclosa bifurca (Araneae: Araneidae) suggests that silk stabilimenta function as camouflage devices. Behaviour, 140(7), 847–868. [Download]


Briceño, R. & Eberhard, W. G. 2002. Decisions during courtship by male and female medflies (Diptera, Tephritidae): correlated changes in male behavior and female acceptance criteria in mass-reared flies. Florida Entomologist, 85(1), 14–31. [Download]
Briceño, R. D. & Eberhard, W. G. 2002. Courtship in the medfly, Ceratitis capitata, includes tactile stimulation with the male’s aristae. Entomologia experimentalis et applicata, 102(3), 221–228. [Download]
Briceño, R. D., Eberhard, W. G., Vilardi, J. C., Liedo, P. & Shelly, T. 2002. Variation in the intermittent buzzing songs of male medflies (Diptera: Tephritidae) associated with geography, mass-rearing, and courtship success. Florida Entomologist, 85(1), 32–40. [Download]
Eberhard, W. G. & Flores, C. 2002. The behavior and natural history of Hybosciara gigantea (Diptera: Sciaridae). Journal of the Kansas Entomological Society, 75(1), 8–15. [Download]
Eberhard, W. G. 2002. Natural history and behavior of Chymomyza mycopelates and C. exophthalma (Diptera: Drosophilidae), and allometry of structures used as signals, weapons, and spore collectors. The Canadian Entomologist, 134(5), 667–687. [Download]
Eberhard, W. G. 2002. Physical restraint or stimulation? The function (s) of the modified front legs of male Archisepsis diversiformis (Diptera, Sepsidae). Journal of Insect behavior, 15(6), 831–850. [Download]
Eberhard, W. G. 2002. The function of female resistance behavior: intromission by male coercion vs. female cooperation in sepsid flies (Diptera: Sepsidae). Revista de Biologia Tropical, 50(2), 485–505. [Download]
Eberhard, W. G. 2002. The relation between aggressive and sexual behavior and allometry in Palaeosepsis dentatiformis flies (Diptera: Sepsidae). Journal of the Kansas Entomological Society, 75(4), 317–332. [Download]


Eberhard, W. G. 2001. Courtship and multi-stage transfer of material to the female’s wings during copulation in Microsepsis armillata (Diptera: Sepsidae). Journal of the Kansas Entomological Society, 74(2), 70–78. [Download]
Eberhard, W. G. 2001. Genitalic behavior in Hybosciara gigantea (Diptera: Sciaridae) and the evolution of species-specific genitalia. Journal of the Kansas Entomological Society, 74(1), 1–9. [Download]
Eberhard, W. G. 2001. Multiple origins of a major novelty: moveable abdominal lobes in male sepsid flies (Diptera: Sepsidae), and the question of developmental constraints. Evolution & Development, 3(3), 206–222. [Download]
Eberhard, W. G. 2001. Species-specific genitalic copulatory courtship in sepsid flies (Diptera, Sepsidae, Microsepsis) and theories of genitalic evolution. Evolution, 55(1), 93–102. [Download]
Eberhard, W. G. 2001. The functional morphology of species-specific clasping structures on the front legs of male Archisepsis and Palaeosepsis flies (Diptera, Sepsidae). Zoological Journal of the Linnean Society, 133(3), 335–368. [Download]
Eberhard, W. G. 2001. Trolling for water striders: active searching for prey and the evolution of reduced webs in the spider Wendilgarda sp.(Araneae, Theridiosomatidae). Journal of Natural History, 35(2), 229–251. [Download]
Eberhard, W. G. 2001. Under the influence: Webs and building behavior of Plesiometa argyra (Araneae, Tetragnathidae) when parasitized by Hymenoepimecis argyraphaga (Hymenoptera, Ichneumonidae). Journal of Arachnology, 29(3), 354–366. [Download]


Briceño, R. D. & Eberhard, W. G. 2000. Male wing positions during courtship by Mediterranean fruit flies (Ceratitis capitata) (Diptera: Tephritidae). Journal of the Kansas Entomological Society, 73(3), 143–147. [Download]
Eberhard, W. G. & Garcia-C, J. M. 2000. Ritual jousting by horned Parisoschoenus expositus weevils (Coleoptera, Curculionidae, Baridinae). Psyche, 103(1-2), 55–84. [Download]
Eberhard, W. G. 2000. Behavior and reproductive status of Microsepsis armillata (Diptera: Sepsidae) flies away from oviposition sites. Annals of the Entomological Society of America, 93(4), 966–971. [Download]
Eberhard, W. G. 2000. Breaking the mold: behavioral variation and evolutionary innovation in Wendilgarda spiders (Araneae Theridiosomatidae). Ethology Ecology & Evolution, 12(3), 223–235. [Download]
Eberhard, W. G. 2000. Criteria for demonstrating postcopulatory female choice. Evolution, 54(3), 1047–1050. [Download]
Eberhard, W. G. 2000. Manipulación de una araña por la larva de una avispa. Una avispa parásita obliga a su hospedera a tejer una tela especial para sus propios fines. Nature. 406 [Download]
Eberhard, W. G. 2000. Spider manipulation by a wasp larva. Nature, 406 (6793), 255–256. [Download]
Eberhard, W. G. 2000. The natural history and behavior of Hymenoepimecis argyraphaga (Hymenoptera: Ichneumonidae) a parasitoid of Plesiometa argyra (Araneae: Tetragnathidae). Journal of Hymenoptera Research, 9(2), 220–240. [Download]
Eberhard, W. G., Garcia-C, M. & Lobo, J. 2000. Size–specific defensive structures in a horned weevil confirm a classic battle plan: avoid fights with larger opponents. Proceedings of the Royal Society of London. Series B: Biological Sciences, 267(1448), 1129-1134. [Download]


Briceño, R. D., Ramos, D. & Eberhard, W. G. 1999. Aggressive behaviour in medflies (Ceratitis capitata) and the effects of mass rearing (Diptera: Tephritidae). Journal of the Kansas Entomological Society, 72(1), 17–27. [Download]
Eberhard, W. G. 1999. Mating systems of sepsid Flies and sexual behavior away from oviposition sites by Sepsis neocynipsea (Diptera: Sepsidae). Journal of the Kansas Entomological Society, 72(1), 129-130. [Download]
Eberhard, W. G. 1999. Mortal mimicry in the fly, Plocoscelus sp. (Diptera, Micropezidae). Biotropica, 31(3), 535–535. [Download]
Eberhard, W. G., Huber, B. & Rodriguez, R. 1999. Don’t forget the biology: a reply to Green. Evolution, 53(5), 1624–1627. [Download]


Briceño, R. D. & Eberhard, W. G. 1998. Medfly courtship duration: a sexually selected reaction norm changed by crowding. Ethology Ecology and Evolution, 10(4), 369–382. [Download]
Eberhard, W. G. & Huber, B. A. 1998. Copulation and sperm transfer in Archisepsis flies (Diptera: Sepsidae) and the evolution of their intromittent genitalia. Studia dipterologica, 5(2), 217–248. [Download]
Eberhard, W. G. & Huber, B. A. 1998. Courtship, copulation, and sperm transfer in Leucauge mariana (Araneae, Tetragnathidae) with implications for higher classification. Journal of Arachnology, 26(3), 342–368. [Download]
Eberhard, W. G. & Huber, B. A. 1998. Possible links between embryology, lack of innervation, and the evolution of male genitalia in spiders. Bulletin of the British Arachnological Society, 11(2), 73–80. [Download]
Eberhard, W. G. 1998. Female roles in sperm competition. pp. 91-116 in T. Birkhead & A. P. Moller (eds.) Sperm competition and sexual selection. New York: Academic Press. [Download]
Eberhard, W. G. 1998. Importancia de la elección femenina críptica para la etología. Etología., 6, 1–8. [Download]
Eberhard, W. G. 1998. Reproductive Behavior of Glyphidops flavifrons and Nerius plurivitatus (Diptera, Neriidae). Journal of the Kansas Entomological Society, 71(2), 89–107. [Download]
Eberhard, W. G. 1998. Sexual behavior of Acanthocephala declivis guatemalana (Hemiptera: Coreidae) and the allometric scaling of their modified hind legs. Annals of the Entomological Society of America, 91(6), 863–871. [Download]
Eberhard, W. G., Huber, B. A., S, R. L. R., Briceño, R. D., Salas, I. & Rodriguez, V. 1998. One size fits all? Relationships between the size and degree of variation in genitalia and other body parts in twenty species of insects and spiders. Evolution, 52(2), 415–431. [Download]
Mendez, V., Briceño, R. D. & Eberhard, W. G. 1998. Functional significance of the capitate supra-fronto-orbital bristles of male medflies (Ceratitis capitata) (Diptera, Tephritidae). Journal of the Kansas Entomological Society, 71(2), 164–174. [Download]


Eberhard, W. G. 1997. Sexual selection by cryptic female choice in insects and arachnids. pp. 32-57 in J. C. Choe & B. J. Crespi (eds.) The evolution of mating systems in insects and arachnids. New York: Cambridge University Press. [Download]
Huber, B. A. & Eberhard, W. G. 1997. Courtship, copulation, and genital mechanics in Physocyclus globosus (Araneae, Pholcidae). Canadian Journal of Zoology, 75(6), 905–918. [Download]


Briceño, R. D., Ramos, D. & Eberhard, W. G. 1996. Courtship behavior of male Ceratitis capitata (Diptera: Tephritidae) in captivity. Florida Entomologist, 79(2), 130–143. [Download]
Eberhard, W. G. & Kariko, S. J. 1996. Copulation behavior inside and outside the beetle Macrohaltica jamaicensis (Coleoptera, Chrysomelidae). Journal of Ethology, 14(1), 59–72. [Download]
Eberhard, W. G. & Marin, M. C. 1996. Sexual behavior and the enlarged hind legs of male Megalopus armatus (Coleoptera, Chrysomelidae, Megalopodinae). Journal of the Kansas Entomological Society, 69(1), 1–8. [Download]
Eberhard, W. G. & Pereira, F. 1996. Functional morphology of male genitalic surstyli in the dungflies Achisepsis diversiformis and A. ecalcarata (Diptera: Sepsidae). Journal of the Kansas Entomological Society, 69(4 suppl), 43–60. [Download]


Briceño, R. D. & Eberhard, W. G. 1995. The functional morphology of male cerci and associated characters in 13 species of tropical earwigs (Dermaptera: Forficulidae, Labiidae, Carcinophoridae, Pygidicranidae). Smithsonian Contributions to Zoology., 555(I-III), 1–63. [Download]
Eberhard, W. G. & Cordero, C. 1995. Sexual selection by cryptic female choice on male seminal products-a new bridge between sexual selection and reproductive physiology. Trends in Ecology & Evolution, 10(12), 493–496. [Download]
Eberhard, W. G. & Pereira, F. 1995. The process of intromission in the Mediterranean fruit fly, Ceratitis capitata (Diptera: Tephritidiae). Psyche, 102(3-4), 99–120. [Download]
Eberhard, W. G. 1995. The web and building behavior of Synotaxus ecuadorensis (Araneae, Synotaxidae). Journal of Arachnology, 23(1), 25–30. [Download]
Hormiga, G., Eberhard, W. G. & Coddington, J. A. 1995. Web-construction behavior in Australian Phonognatha and the phylogeny of Nephiline and tetragnathid spiders (Araneae, Tetragnathidae). Australian Journal of Zoology, 43(4), 313–364. [Download]
R. D. Briceño, D. Ramos, and William G. Eberhard. 1996. Courtship Behavior of Male Ceratitis capitata (Diptera: Tephritidae) in Captivity. [Download]


Eberhard W. 1994. The encyclopedia of land invertebrate behaviour. The Quarterly Review of Biology.69 (4),559-560. [Download]
Eberhard, W. G. 1994. Evidence for widespread courtship during copulation in 131 species of insects and spiders, and implications for cryptic female choice. Evolution, 48(3), 711–733. [Download]


Eberhard, W. G. & Pereira, F. 1993. Ultrastructure of cribellate silk of nine species in eight families and possible taxonomic implications (Araneae: Amaurobiidae, Deinopidae, Desidae, Dictynidae, Filistatidae, Hypochilidae, Stiphidiidae, Tengellidae). Journal of Arachnology, 21, 161–174. [Download]
Eberhard, W. G. 1993. Evaluating models of sexual selection: genitalia as a test case. The American Naturalist, 142(3), 564–571. [Download]
Eberhard, W. G., Achoy, R., Marin, M. & Ugalde, J. 1993. Natural history and behavior of two species of Macrohaltica (Coleoptera: Chrysomelidae). Psyche, 100(1-2), 93–120. [Download]
Eberhard, W. G., Platnick, N. I. & Schuh, R. T. 1993. Natural history and systematics of arthropod symbionts (Araneae; Hemiptera; Diptera) inhabiting webs of the spider Tengella radiata (Araneae, Tengellidae). American Museum Novitates., 3065, 1–17. [Download]


Eberhard, W. G. 1992. Species isolation, genital mechanics, and the evolution of species-specific genitalia in three species of Macrodactylus beetles (Coleoptera, Scarabeidae, Melolonthinae). Evolution, 46(6), 1774–1783. [Download]
Eberhard, W. G. 1992. Web construction by Modisimus sp.(Araneae, Pholcidae). Journal of Arachnology, 20, 25–34. [Download]


Eberhard, W. G. & Gutierrez, E. E. 1991. Male dimorphisms in beetles and earwigs and the question of developmental constraints. Evolution, 45(1), 18–28. [Download]
Eberhard, W. G. 1991. Chrosiothes tonala (Araneae, Theridiidae): a web-building spider specializing on termites. Psyche, 98(1), 7–20. [Download]


Eberhard, W. G. 1990. Early stages of orb construction by Philoponella vicina, Leucauge mariana, and Nephila clavipes (Araneae, Uloboridae and Tetragnathidae), and their phylogenetic implications. Journal of Arachnology, 18(2), 205–234. [Download]
Eberhard, W. G. 1990. Evolution in bacterial plasmids and levels of selection. Quarterly Review of Biology, 65(1), 3–22. [Download]
Eberhard, W. G. 1990. Function and phylogeny of spider webs. Annual Review of Ecology and Systematics, 21, 341–372. [Download]
Eberhard, W. G. 1990. Imprecision in the behavior of Leptomorphus sp.(Diptera, Mycetophilidae) and the evolutionary origin of new behavior patterns. Journal of Insect Behavior, 3(3), 327–357. [Download]
Santana, M., Eberhard, W. G., Bassey, G., Prestwich, K. N. & Briceño, R. D. 1990. Low predation rates in the field by the tropical spider Tengella radiata (Araneae: Tengellidae). Biotropica, 22(3), 305–309. [Download]


Eberhard, W. G. 1989. Effects of orb web orientation and spider size on prey retention. Bulletin of the British Arachnological Society, 8(2), 45–48. [Download]
Eberhard, W. G. 1989. Niche expansion in the spider Wendilgarda galapagensis (Araneae, Theridiosomatidae) on Cocos Island. Revista de Biología Tropical, 37(2), 163–168. [Download]
Eberhard, W. G. 1989. Use of host aggregations as mating sites by Chrysagria alticophaga (Diptera: Sarcophagidae). Psyche, 96(1-2), 45–48. [Download]
Eberhard, W. G. 1989. Why do bacterial plasmids carry some genes and not others? Plasmid, 21(3), 167–174. [Download]
Wcislo, W. T. & Eberhard, W. G. 1989. Club fights in the weevil Macromerus bicinctus (Coleoptera: Curculionidae). Journal of the Kansas Entomological Society, 62(4), 421–429. [Download]


Eberhard, W. G. 1988. Behavioral flexibility in orb web construction: effects of supplies in different silk glands and spider size and weight. Journal of Arachnology, 16(3), 295–302. [Download]
Eberhard, W. G. 1988. Combing and sticky silk attachment behaviour by cribellate spiders and its taxonomic implications. Bulletin of the British Arachnological Society, 7(8), 247–251. [Download]
Eberhard, W. G. 1988. Group nesting in two species of Euglossa bees (Hymenoptera: Apidae). Journal of the Kansas Entomological Society, 61(4), 406–411. [Download]
Eberhard, W. G. 1988. Memory of distances and directions moved as cues during temporary spiral construction in the spider Leucauge mariana (Araneae: Araneidae). Journal of Insect Behavior, 1(1), 51–66. [Download]
Eberhard, W. G. 1988. Paradoxical post-coupling courtship in Himantigera nigrifemorata (Diptera, Stratiomyidae). Psyche, 95 (1-2), 115–122. [Download]
Wcislo, W. T., West-Eberhard, M. J. & Eberhard, W. G. 1988. Natural history and behavior of a primitively social wasp, Auplopus semialatus, and its parasite, Irenangelus eberhardi (Hymenoptera: Pompilidae). Journal of Insect Behavior, 1(3), 247–260. [Download]


Briceno, R. D. & Eberhard, W. G. 1987. Genetic and environmental effects on wing polymorphisms in two tropical earwigs (Dermaptera: Labiidae). Oecologia, 74(2), 253–255. [Download]
Eberhard, W. G. 1987. Construction behavior of non-orb weaving cribellate spiders and the evolutionary origin of orb webs. Bulletin of the British Arachnological Society, 7(6), 175–178. [Download]
Eberhard, W. G. 1987. Effects of gravity on temporary spiral construction by Leucauge mariana (Araneae: Araneidae). Journal of Ethology, 5(1), 29–36. [Download]
Eberhard, W. G. 1987. How spiders initiate airborne lines. Journal of Arachnology, 15, 1–9. [Download]
Eberhard, W. G. 1987. Hub construction by Leucauge mariana (Araneae, Araneidae). Bulletin of the British Arachnological Society, 7(4), 128–132. [Download]
Eberhard, W. G. 1987. Maternal behavior in Apollophanes punctipes (O. Pickard-Cambridge)(Araneae, Thomisidae). Journal of Arachnology, 14, 398. [Download]
Eberhard, W. G. 1987. Runaway sexual selection. Natural History, 96(12), 4-8. [Download]
Eberhard, W. G. 1987. Use of horns in flights by the dimorphic males of Ageopsis nigricollis (Coleoptera, Scarabeidae, Dynastinae). Journal of the Kansas Entomological Society, 60(4), 504-509. [Download]
Eberhard, W. G. 1987. Web-building behavior of anapid, symphytognathid and mysmenid spiders (Araneae). Journal of Arachnology, 14, 339–356. [Download]


Eberhard, W. G. 1986. Behavioral ecology of the tropical damselfly Hetaerina macropus Selys (Zygoptera: Calopterygidae). Odonatologica, 15(1), 51–60. [Download]
Eberhard, W. G. 1986. Effects of orb-web geometry on prey interception and retention. Pp 70-100 in W. A. Shear (ed.) Spiders Webs, Behaviour and Evolution. Palo Alto: Stanford University Press. [Download]
Eberhard, W. G. 1986. Ontogenetic changes in the web of Epeirotypus sp.(Araneae, Theridiosomatidae). Journal of Arachnology, 14(1), 125–128. [Download]
Eberhard, W. G. 1986. Possible mutualism between females of the subsocial membracid Polyglypta dispar (Homoptera). Behavioral Ecology and Sociobiology, 19(6), 447–453. [Download]
Eberhard, W. G. 1986. Pupation in mycetophilid flies: a correction. Psyche, 93(1-2), 117–120. [Download]
Eberhard, W. G. 1986. Subsocial behaviour in the spitting spider Scytodes intricata (Araneae: Scytodidae). Revue Arachnologique 7(1), 35–40. [Download]
Eberhard, W. G. 1986. Trail line manipulation as a character for higher level spider taxonomy. Pp. 49-51 in W. G. Eberhard, Y. D. Lubin & B. C. Robinson (eds.) Proceedings of the Ninth International Congress of Arachnology, Panamá 1983. Washington D.C.: Smithsonian Institution Press. [Download]
Eberhard, W. G. 1986. Why are genitalia good species characters? Proceedings of the Ninth International Congress of Arachnology, Panamá 1983 (William G. Eberhard, Yael D. Lubin, Barbara C. Robinson., pp. 53-59). Washington, D.C. [Download]
Philippi, T. & Eberhard, W. G. 1986. Foraging behavior of Stictia signata (Hymenoptera: Sphecidae). Journal of the Kansas Entomological Society 59(4), 604–608. [Download]


Eberhard, W. G. & Briceño, R. D. 1985. Behavior and ecology of four species of Modisimus and Blechroscelis (Araneae, Pholcidae). Revue Arachnologique, 6(1), 29–36. [Download]
Eberhard, W. G. 1985. The «sawtoothed» orb web of Eustala sp.(Araneae, Araneidae), with a discussion of ontogenetic changes in spiders’ web-building behavior. Psyche, 92(1), 105–118. [Download]


Opell, B. D. & Eberhard, W. G. 1984. Resting postures of orb-weaving uloborid spiders (Araneae, Uloboridae). Journal of Arachnology, 11, 369–376. [Download]


Castillo, J. & Eberhard, W. G. 1983. Use of artificial webs to determine prey available to orb weaving spiders. Ecology, 64(6), 1655–1658. [Download]
Eberhard, W. G. 1983. Behavior of adult bottle brush weevils (Rhinostomus barbirostris)(Coleoptera: Curculionidae). Revista de Biologia Tropical, 31(2), 233–244. [Download]
Eberhard, W. G. 1983. Predatory behaviour of an assassin spider, Chorizopes sp.(Araneidae), and the defensive behavior of its prey. Journal of the Bombay Natural History Society, 79(3), 522–524. [Download]


Eberhard, W. G. 1982. Beetle horn dimorphism: making the best of a bad lot. The American Naturalist, 119(3), 420–426. [Download]
Eberhard, W. G. 1982. Behavioral characters for the higher classification of orb-weaving spiders. Evolution, 36(5), 1067–1095. [Download]
Eberhard, W. G. 1982. Review: Behavior of Arachnids. Science, New Series, 216(4548), 876-877. [Download]
Lubin, Y. D., Opell, B. D., Eberhard, W. G. & Levi, H. W. 1982. Orb plus cone-webs in Uloboridae (Araneae), with a description of a new genus and four new species. Psyche, 89(1-2), 29–64. [Download]


Eberhard, W. G. 1981. Construction behaviour and the distribution of tensions in orb webs. Bulletin of the British Arachnological Society, 5(5), 189–204. [Download]
Eberhard, W. G. 1981. Notes on the natural history of Taczanowskia sp.(Araneae: Araneidae). Bulletin of the British Arachnological Society, 5(4), 175–176. [Download]
Eberhard, W. G. 1981. The natural history of Doryphora sp.(Coleoptera, Chrysomelidae) and the function of its sternal horn. Annals of the Entomological Society of America, 74(5), 445–448. [Download]
Eberhard, W. G. 1981. The single line web of Phoroncidia studo Levi (Araneae, Theridiidae): a prey attractant? Journal of Arachnology, 9, 229–232. [Download]


Chacón, P. & Eberhard, W. G. 1980. Factors affecting numbers and kinds of prey caught in artificial spider webs, with considerations of how orb webs trap prey. Bulletin of the British Arachnological Society 5(1): 29-38. [Download]
Eberhard, W. G. 1980. Evolutionary consequences of intracellular organelle competition. The Quarterly Review of Biology, 55(3), 231-249. [Download]
Eberhard, W. G. 1980. Persistent stickiness of cribellum silk. Journal of Arachnology, 8(3), 283. [Download]
Eberhard, W. G. 1980. Spider and fly play cat and mouse. Natural History, 56-61. [Download]
Eberhard, W. G. 1980. The natural history and behavior of the bolas spider, Mastophora dizzydeani sp. n.(Araneae). Psyche, 87(3-4), 143–170. [Download]


Eberhard, W. G. 1979. Argyrodes attenuatus (Theridiidae): a web that is not a snare. Psyche, 86(4), 407–414. [Download]
Eberhard, W. G. 1979. Rates of egg production by tropical spiders in the field. Biotropica, 11(4), 292–300. [Download]
Eberhard, W. G. 1979. The function of horns in Podischnus agenor (Dynastinae) and other beetles. Sexual selection and reproductive competition in insects, 463, 231-258. [Download]
Lahmann-Zeledón, E. J. & Eberhard, W. G. 1979. Factores selectivos que afectan la tendencia a agruparse en la araña colonial Philoponella semiplumosa (Aranae; Uloboridae). Revista de Biología Tropical., 27(2), 231–240. [Download]


Eberhard, W. G. 1978. Mating swarms of a South American Acropygia (Hymenoptera: Formicidae). Entomological News, 89(1/2), 14–16. [Download]
Eberhard, W. G., Barreto, M. & Pfizenmaier, W. 1978. Web robbery by mature male orb-weaving spiders. Bulletin of the British Arachnological Society, 4(5), 228–230. [Download]
Lubin, Y. D., Eberhard, W. G. & Montgomery, G. B. 1978. Webs of Miagrammopes (Araneae: Uloboridae) in the neotropics. Psyche, 85(1), 1–23. [Download]


Eberhard, W. G. 1977. Aggressive chemical mimicry by a bolas spider. Science, 198(4322), 1173-1175. [Download]
Eberhard, W. G. 1977. Artificial spider webs. Bulletin of the British Arachnological Society, 4(3), 126–128. [Download]
Eberhard, W. G. 1977. Fighting behavior of male Golofa porteri beetles (Scarabeidae: Dynastinae). Psyche, 84(3-4), 292–298. [Download]
Eberhard, W. G. 1977. La ecología y comportamiento de los adultos del cucarrón Podischnus agenor. Revista Colombiana de Entomología, 3(1-2), 17–21. [Download]
Eberhard, W. G. 1977. The webs of newly emerged Uloborus diversus and of a male Uloborus sp.(Araneae: Uloboridae). Journal of Arachnology, 4, 201–206. [Download]
Eberhard, W. G. 1977. ‘Rectangular orb’ webs of Synotaxus (Araneae: Theridiidae). Journal of Natural History, 11(5), 501-507. doi:10.1080/00222937700770431 [Download]
William G. Eberhard. 1977. Mimetismo Químico agresivo de una araña Bolera. Science Vol. 198. 122-124 [Download]


Eberhard, W. G. 1976. Photography of orb webs in the field. Bulletin of the British Arachnological Society, 3(7), 200–204. [Download]
Eberhard, W. G. 1976. Physical properties of sticky spirals and their connections: sliding connections in orb webs. Journal of Natural History, 10(5), 481–488. [Download]


Eberhard, W. G. 1975. The ecology and behavior of a subsocial pentatomid bug and two scelionid wasps: strategy and counterstrategy in a host and its parasites. Smithsonian Contributions to Zoology, 205, 1-39. [Download]
Eberhard, W. G. 1975. The ‘inverted ladder’orb web of Scoloderus sp. and the intermediate orb of Eustala (?) sp. Araneae: Araneidae. Journal of Natural History, 9(1), 93–106. [Download]


Eberhard, W. G. 1974. Maternal behavior in a South American Lyssomanes. Bulletin of the British Arachnological Society, 3(2), 51. [Download]
Eberhard, W. G. 1974. The natural history and behaviour of the wasp Trigonopsis cameronii Kohl (Sphecidae). Transactions of the Royal Entomological Society of London, 125(3), 295–328. doi:10.1111/j.1365-2311.1973.tb00545.x [Download]


Eberhard, W. G. 1973. Stabilimenta on the webs of Uloborus diversus (Araneae: Uloboridae) and other spiders. Journal of Zoology, 171(3), 367–384. [Download]
Eberhard, W. G. 1973. The ability of a spitting spider, Scytodes hebraica, to capture flying prey. Entomological News, 84, 156. [Download]


Eberhard, W. G. 1972. Altruistic behavior in a sphecid wasp: support for kin-selection theory. Science, 175(4028), 1390-1391. [Download]
Eberhard, W. G. 1972. Observations on the biology of Achaearanea tesselata (Araneae: Theridiidae). Psyche, 79(3), 209–212. [Download]
Eberhard, W. G. 1972. The web of Uloborus diversus (Araneae: Uloboridae). Journal of Zoology, 166(4), 417–465. [Download]


Eberhard, W. G. 1971. Senile web patterns in Uloborus diversus (Araneae: Uloboridae). Developmental Psychobiology, 4(3), 249–254. doi:10.1002/dev.420040303 [Download]
Eberhard, W. G. 1971. The ecology of the web of Uloborus diversus (Araneae: Uloboridae). Oecologia, 6(4), 328–342. [Download]
Eberhard, W. G. 1971. The Predatory Behavior of Two Wasps, Agenoideus humilis (Pompilidae) and Sceliphron caementarium (Sphecidae), on the Orb Weaving Spider, Araneus cornutus (Araneidae). Psyche 77:243-251. [Download]


Eberhard, W. G. 1970. The natural history of the fungus gnats Leptomorphus bifasciatus (Say) and L. subcaeruleus (Coquillett)(Diptera: Mycetophilidae). Psyche, 77(3), 361–384. [Download]
Eberhard, W. G. 1970. The predatory behavior of two wasps, Agenoideus humilis (Pompilidae) and Sceliphron caementarium (Sphecidae), on the orb weaving spider, Araneus cornutus (Araneidae). Psyche, 77(2), 243–251. [Download]


Eberhard W. 1969. The Predatory Behavior of Two Wasps, Agenoideus humilis (Pompilidae) and Sceliphron caementarium (Sphecidae), on the Orb Weaving Spider, Araneus cornutus (Araneidae). Psyche 77:243-251. [Download]
Eberhard, W. G. & Levi, H. W. 1969. Review: Silk Making. Science 164(3882), 941-942. [Download]
Eberhard, W. G. 1969. Computer simulation of orb-web construction. American Zoologist, 9(1), 229. [Download]
Langer, R. M. & Eberhard, W. G. 1969. Laboratory photography of spider silk. American Zoologist, 9(1), 97. [Download]


Eberhard, W. G. 1967. Attack behavior of diguetid spiders and the origin of prey wrapping in spiders. Psyche, 74(2), 173–181. [Download]